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A character has undergone a complete physical sex change, usually through magic or Applied Phlebotinum.
Depending on the medium, genre, and storyline, this may be a one-time temporary change, a recurring change (causing the character to jump the gender line often), or even permanent.
If a man and a woman bodies, it's a variant of "Freaky Friday" Flip.
For this trope's impact on relationships, see Unsettling Gender Reveal, Jumping the Gender Barrier and Gender-Bender Friendship.
Contrast with Transsexual and Hermaphrodite, which are real-life phenomena (though on rare occasions, a Gender Bender character might a hermaphrodite instead of, strictly speaking, the "opposite sex"; also, Gender Bender stories can intersect with transsexual characters, by having someone transition unexpectedly by magic, or have the experience of being gender-changed bring a transsexual character out of denial.
Not to mention all the possibilities of using Gender Bender stories as metaphors for transsexuality...).
Also contrast with Easy Sex Change, which deals with idealized but realistic takes on the Transsexual experience. If the character belongs to a species that does this as regular thing, see Sex Shifter.